Ray Romano Manny (voice) John Leguizamo Sid (voice) Denis Leary Diego (voice) Goran Visnjic Soto (voice) Jack Black Zeke (voice) Cedric the Entertainer Carl (voice) Stephen Root Frank / Start (voice) Diedrich Bader Oscar (voice) Alan Tudyk Lenny / Oscar / Dab (voice) Lorri Bagley Jennifer (voice) Jane Krakowski Rachel (voice) Josh Hamilton Dodo / Aardvark (voice) Chris Wedge Dodo / Scrat (voice) Denny Dillon Glyptodon (voice) Mitzi McCall Glyptodont (voice) Tara Strong Baby Moeritherium (voice) Darin De Paul Additional Voices (voice) (uncredited) Dann Fink Dodo (voice) (uncredited) Patrick Pinney Various Mammals (uncredited) Phil Proctor Various Mammals (uncredited) Ray Romano Manny (voice) John Leguizamo Sid (voice) Denis Leary Diego (voice) Goran Visnjic Soto (voice) Jack Black Zeke (voice) Cedric the Entertainer Carl (voice) Stephen Root Frank / Start (voice) Diedrich Bader Oscar (voice) Alan Tudyk Lenny / Oscar / Dab (voice) Lorri Bagley Jennifer (voice) Jane Krakowski Rachel (voice) Josh Hamilton Dodo / Aardvark (voice) Chris Wedge Dodo / Scrat (voice) Denny Dillon Glyptodon (voice) Mitzi McCall Glyptodont (voice) Tara Strong Baby Moeritherium (voice) Darin De Paul Additional Voices (voice) (uncredited) Dann Fink Dodo (voice) (uncredited) Patrick Pinney Various Mammals (uncredited) Phil Proctor Various Mammals (uncredited)